Peer-reviewed papers

For citations and metrics, please see my Google Scholar profile. Also, see book chapters, editorials, and my popular science articles.

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Preprints (not yet peer-reviewed)

Stervander M, Melo M, Jones P & Hansson B. 2022. Genomic signatures of isolation, hybridization, and selection during speciation of island finches. bioRxiv 2022.01.05.474904. DOI: 10.1101/2022.01.05.474904.

Read the non-reviewed preprint manuscript at bioRxiv.
Datasets freely available at Zenodo: DOI



Brown AF, Lawrie Y, Shannon TJ, Collinson JM, Kirwan GM, Kirkconnell A & Stervander M. 2022. First genetic data for the Critically Endangered Cuban endemic Zapata Rail Cyanolimnas cerverai, and the taxonomic implications. Journal of Ornithology, in press. DOI: 10.1007/s10336-022-02004-0.

Read Open Access paper at Journal of Ornithology.

The non-reviewed preprint manuscript prior to revisions is available at bioRxiv.
Datasets freely available at Zenodo: DOI


Roved J, Hansson B, Stervander M, Hasselquist D* & Westerdahl H*. 2022. MHCtools – an R package for MHC high-throughput sequencing data: genotyping, haplotype and supertype inference, and downstream genetic analyses in non-model organisms. Molecular Ecology Resources 22: 2775–2792. DOI: 10.1111/1755-0998.13645. *Shared last authorship.

Read Open Access paper at Molecular Ecology Resources.

The non-reviewed preprint manuscript prior to revisions is available at bioRxiv.
Datasets freely available at Zenodo: DOI


Stervander M & Cresko WA. 2021. A highly contiguous nuclear genome assembly of the mandarinfish Synchiropus splendidus (Syngnathifromes: Callionymidae). G3: Genes | Genomes | Genetics 11: jkab306. DOI: 10.1093/g3journal/jkab306.

Read Open Access paper at G3: Genes | Genomes | Genetics.


Featured by The Scientist.


Stervander M & Haelewaters D. 2020. Fishiness of piscine birds linked to absence of poisonous fungi but not pizza. Oceanography & Fisheries Open Access Journal 12(5): 555850. DOI: 10.19080/OFOAJ.2020.12.555850.

Read this ploy Open Access “paper” at Oceanography & Fisheries Open Access Journal, and marvel at the astonishing scientific breakthroughs that we present. Or perhaps be bewildered over the predatory journal industry publishing our high-quality gobbledegook.


Stervander M, Hansson B, Olsson U, Hulme MF, Ottosson U & Alström P. 2020. Molecular species delimitation of larks (Aves: Alaudidae), and integrative taxonomy of the genus Calandrella, with the description of a range-restricted African relic taxon. Diversity 12: 428. DOI: 10.3390/d12110428.

Read Open Access paper at Diversity.
Datasets freely available at Zenodo: DOI


Chaves JA, Martinez-Torres PJ, Depino EA, Espinoza-Ulloa S, García-Loor J, Beichman A & Stervander M. 2020. Evolutionary history of the Galápagos Rail revealed by ancient mitogenomes and modern samples. Diversity 12: 425. DOI: 10.3390/d12110425.

Read Open Access paper at Diversity.
Datasets freely available at Zenodo: DOI
The non-reviewed preprint manuscript prior to revisions is available at bioRxiv.

The cover of Diversity volume 12 issue 11, featuring Chaves et al. (2020)

Feng S*, Stiller J*, Deng Y*, Armstrong J*, Fang Q, Reeve AH, Xie D, Chen G, Guo C, Faircloth BC, Petersen B, Wang Z, Zhou Q, Diekhans M, Chen W, Andreu-Sánchez S, Margaryan A, Howard JT, Parent C, Pacheco G, Sinding MHS, Puetz L, Cavill E, Ribeiro ÂM, Eckhart L, Fjeldså J, Hosner P, Brumfield RT, Christidis L, Bertelsen MF, Sicheritz-Ponten T, Tietze DT, Robertson BC, Song G, Borgia G, Claramunt S, Lovette IJ, Cowen SJ, Njoroge P, Dumbacher JP, Ryder OA, Fuchs J, Bunce M, Burt DW, Cracraft J, Meng G, Hackett SJ, Ryan PG, Jønsson KA, Jamieson IG, da Fonseca RR, Braun EL, Houde P, Mirarab S, Suh S, Hansson B, Ponnikas S, Sigeman S, Stervander M, Frandsen PB, van der Zwan H, van der Sluis R, Visser C, Balakrishnan CN, Clark AG, Fitzpatrick JW, Bowman R, Chen N, Cloutier A, Sackton TB, Edwards S, Foote DJ, Shakya SB, Sheldon SH, Vignal A, Soares AER, Shapiro B, González-Solís J, Ferrer-Obiol J, Rozas J, Riutort R, Tigano A, Friesen V, Dalén L, Urrutia AO, Székely T, Liu Y, Campana MG, Corvelo A, Fleischer RC, Rutherford KM, Gemmell NJ, Dussex N, Mouritsen H, Thiele N, Delmore K, Liedvogel M, Franke A, Hoeppner MP, Krone O, Fudickar AM, Milá B, Ketterson ED, Fidler AE, Friis G, Parody-Merino ÁM, Battley PF, Cox MP, Lima NCB, Prosdocimi F, Parchman TL, Schlinger BA, Loiselle BA, Blake JG, Lim HC, Day LB, Fuxjager MJ, Baldwin MW, Braun MJ, Wirthlin M, Dikow RB, Ryder TB, Camenisch G, Keller LF, DaCosta JM, Hauber ME, Louder MIM, Witt CC, McGuire JA, Mudge J, Megna LC, Carling MD, Wang B, Taylor SA, Del-Rio G, Aleixo A, Vasconcelos ATR, Mello CV, Weir JT, Haussler D, Li Q, Yang H, Wang J, Lei F, Rahbek C, Gilbert MTP, Graves GR, Jarvis ED, Paten B & Zhang G. 2020. Dense sampling of bird diversity increases power of comparative genomics. Nature 587: 252–257. DOI: 10.1038/s41586-020-2873-9. *Shared first authorship.

Read Open Access paper at Nature.

The cover of Nature volume 587 issue 7833, featuring Feng, Stiller, Deng, Armstrong et al. (2020)

Stervander M*, Dierickx EG*, Thorley J*, Brooke M de L & Westerdahl H. 2020. High MHC gene copy number maintains diversity despite homozygosity in a Critically Endangered single-island endemic bird, but no evidence of MHC-based mate choice. Molecular Ecology 29: 3578–3592. DOI: 10.1111/mec.15471. *Shared first authorship.

Read Open Access paper at Molecular Ecology. The non-reviewed preprint manuscript prior to revisions is available at bioRxiv.
Datasets freely available at Zenodo: DOI

Selected as “From the Cover” paper. Read the News and Views perspective about the paper.

The cover of Molecular Ecology volume 29, issue 19, featuring Stervander et al. (2020)

Gillespie RG, Bennett GM, De Meester L, Feder JL, Fleischer RC, Harmon LJ, Hendry AP, Knope M, Mallet J, Martin C, Parent CE, Patton AH, Pfennig KS, Rubinoff D, Schluter D, Seehausen O, Shaw KL, Stacy E, Stervander M, Stroud JT, Wagner C & Wogan GOU. 2020. Comparing adaptive radiations across space, time, and taxa. Journal of Heredity 111: 1–20. DOI: 10.1093/jhered/esz064.

Read Open Access paper at Journal of Heredity.

The cover of Journal of Heredity volume 111, issue 1, featuring Gillespie et al. (2020)


Lehikoinen A, Lindén A, Karlsson M, Andersson A, Crewe TL, Dunn EH, Gregory G, Karlsson L, Kristiansen V, Mackenzie S, Newman S, Røer JE, Sharpe C, Sokolov LV, Steinholtz Å, Stervander M, Takats Priestley L, Tirri I-S & Skjold Tjørnløv R. 2019. Phenology of the avian spring migratory passage in Europe and North America: asymmetric advancement in time and increase in duration. Ecological Indicators 101: 985–991. DOI: 10.1016/j.ecolind.2019.01.083.

Read paper at Ecological Indicators.


Inaccessible Rail (photo: Peter G. Ryan) Stervander M, Ryan PG, Melo M & Hansson B. 2019. The origin of the world’s smallest flightless bird, the Inaccessible Island Rail Atlantisia rogersi (Aves: Rallidae). Molecular Phylogenetics and Evolution 130: 92–98. DOI: 10.1016/j.ympev.2018.10.007.

Read paper at Molecular Phylogenetics and Evolution. Datasets freely available at Zenodo: DOI

Media coverage from 200+ outlets in 30+ languages, with links to articles in e.g. Smithsonian Magazine, Atlas Obscura, and Gizmodo.


Hansson2018MER Hansson B, Sigeman H*, Stervander M*, Tarka M, Ponnikas S, Strandh M, Westerdahl H & Hasselquist D. 2018. Contrasting results from GWAS and QTL mapping on wing length in great reed warblers. Molecular Ecology Resources 18: 867–876. DOI: 10.1111/1755-0998.12785. *Equal contribution.
Read paper at Molecular Ecology Resources.


Stervander2017OS_nitidusStervander M, Knudsen H & Bruun Kristensen A. 2017. Green Warbler Phylloscopus nitidus ringed at Blåvand: molecular confirmation of a Danish first and European eighth vagrant record. Ornis Svecica 27: 121–131. DOI: 10.34080/os.v27.19565.

Read Open Access paper at Ornis Svecica.


Melo2017Ibis_bMelo M*, Stervander M*, Hansson B & Jones P. 2017. The endangered São Tomé Grosbeak Neospiza concolor is the world’s largest canary. Ibis 159: 673–679. DOI: 10.1111/ibi.12466. *Shared first authorship.

Read paper at Ibis. Read my popular summary blog post and our post on the BOU blog. Media coverage from 60 outlets in four languages including e.g. National Geographic and New Scientist.



Stervander2016MPEStervander M, Alström P, Olsson U, Ottosson U, Hansson B & Bensch S. 2016. Multiple instances of paraphyletic species and cryptic taxa revealed by mitochondrial and nuclear RAD data for Calandrella larks (Aves: Alaudidae). Molecular Phylogenetics and Evolution 102: 233–245. DOI: 10.1016/j.ympev.2016.05.032.

Read paper at MPE. Data freely available at DataDryad. Read popular summary blog post.


Stervander2015MolEcol_CterStervander M, Illera JC, Kvist L, Barbosa P°, Keehnen NP°, Pruisscher P°, Bensch S & Hansson B. 2015. Disentangling the complex evolutionary history of the Western Palearctic blue tits (Cyanistes spp.) – phylogenomic analyses suggest radiation by multiple colonization events and subsequent isolation. Molecular Ecology 24(10): 2477–2494. DOI: 10.1111/mec.13145. °Mentored undergraduate student.

Data freely available at DataDryad. Read paper at Molecular Ecology. See blog post and discussion


Sjögren2013Sjögren J, Struwe W, Cosgrave E, Rudd P, Stervander M, Allhorn M, Hollands A, Nizet V & Collin M. 2013. EndoS2 is a unique and conserved enzyme of serotype M49 group A Streptococcus that hydrolyses N-linked glycans on IgG and α1-acid glycoprotein. Biochemical Journal 455: 107–118. DOI: 10.1042/BJ20130126.

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Iwajomo2013Iwajomo SB, Stervander M, Helseth A & Ottosson U. 2013. Population trends and migration strategy of the Wood Sandpiper Tringa glareola at Ottenby, SE Sweden. Ringing & Migration 28(1): 6–15. DOI: 10.1080/03078698.2013.811157.

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Hellgren2013_aHellgren O, Wood MJ, Waldenström J, Hasselquist D, Ottosson U, Stervander M & Bensch S. 2013. Circannual variation in blood parasitism in a sub-Saharan migrant, the garden warbler. Journal of Evolutionary Biology 26(5): 1047–1059. DOI: 10.1111/jeb.12129.

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Pala2011Pala I, Naurin S, Stervander M, Hasselquist D, Bensch S & Hansson B. 2011. Evidence of a neo-sex chromosome in birds. Heredity 108(3): 264–272. DOI: 10.1038/hdy.2011.70.

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Jonzén N, Lindén A, Ergon T, Knudsen E, Vik JO, Rubolini D, Piacentini D, Brinch C, Spina F, Karlsson L, Stervander M, Andersson A, Waldenström J, Lehikoinen A, Edvardsen E, Solvang R & Stenseth NC. 2007. Response to comment on “Rapid advance of spring arrival dates in long-distance migratory birds”. Science 315: 598c. DOI: 10.1126/science.1136920. Undergraduate author.

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Wallensten2011EIDWallensten A, Munster VJ, La Torre-Magalef N, Brytting M, Elmberg J, Fouchier RAM, Fransson T, Haemig PD, Karlsson M, Lundkvist Å, Osterhaus ADME, Stervander M, Waldenström J, & Olsen B. 2007. Surveillance of Influenza A Virus in migratory waterfowl in northern Europe. Emerging Infectious Diseases 13(3): 404–411. DOI: 10.3201/eid1303.061130. Undergraduate author.

Read paper. Highly cited


Mallards in Castle Park, by postman.peter (CC BY 2.0). Click image for details.Wallensten A, Munster VJ, Karlsson M, Lundkvist Å, Brytting M, Stervander M, Osterhaus ADME, Fouchier RAM & Olsen B. 2006. High prevalence of influenza A virus in ducks caught during spring migration through Sweden. Vaccine 24: 6734–6735. DOI: 10.1016/j.vaccine.2006.05.057. Undergraduate author.

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Jonzen2006ScienceJonzén N, Lindén A, Ergon T, Knudsen E, Vik JO, Rubolini D, Piacentini D, Brinch C, Spina F, Karlsson L, Stervander M, Andersson, A, Waldenström J, Lehikoinen A, Edvardsen E, Solvang R & Stenseth NC. 2006. Rapid advance of spring arrival dates in long-distance migratory birds. Science 312: 1959–1961. DOI: 10.1126/science.1126119. Undergraduate author.

Read paper. Highly cited. 2006 Classic Paper according to Google Scholar. Featured in the book Science Magazine’s State of the Planet 2008–2009.


Jonzen2006OSJonzén N, Piacentini D, Andersson A, Montemaggiori A, Stervander M, Rubolini D, Waldenström J & Spina F. 2006. The timing of spring migration in trans-Saharan migrants: a comparison between Ottenby, Sweden, and the island of Capri, Italy. Ornis Svecica 16: 27–33. DOI: 10.34080/os.v16.22412. Undergraduate author.

Read Open Access paper at Ornis Svecica.


Helseth2005Ardea_b_knotHelseth A, Lindström Å & Stervander M. 2005. Southward migration and fuel deposition of Red Knots Calidris canutus. Ardea 93(2): 213–224. Authored before entering university.

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Little rush warbler by Alan Manson (CC BY-SA 2.0). Click image for details.Stervander M†✉, Ottosson U, Hulme M & Molokwu MN. 2005. Little Rush Warbler Bradypterus baboecala new to Plateau state, Nigeria. Malimbus 27: 46–47. Authored before entering university.

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Helseth2005OS_aHelseth A, Stervander M & Waldenström J. 2005. Migration patterns, population trends and morphometrics of the Ruddy Turnstone Arenaria interpres passing Ottenby in south-eastern Sweden. Ornis Svecica 15: 63–72. DOI: 10.34080/os.v15.22411. Authored before entering university.

Read Open Access paper at Ornis Svecica or see updated pdf.


Stervander2005JABStervander M†✉, Lindström Å, Jonzén N & Andersson A. 2005. Timing of spring migration in birds: long-term trends, North Atlantic Oscillation and the significance of different migration routes. Journal of Avian Biology 36: 210–221. DOI: 10.1111/j.0908-8857.2005.03360.x. Authored before entering university.

Read paper. Highly cited.